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  1. Robert William Holley (January 28, 1922 – February 11, 1993) was an American biochemist. He shared the Nobel Prize in Physiology or Medicine in 1968 (with Har Gobind Khorana and Marshall Warren Nirenberg) for describing the structure of alanine transfer RNA, linking DNA and protein synthesis .

  2. Robert William Holley was an American biochemist who shared the Nobel Prize in Physiology or Medicine in 1968 with Marshall Warren Nirenberg and Har Gobind Khorana. Their research helped explain how the genetic code controls the synthesis of proteins. Holley obtained his Ph.D. in organic chemistry.

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  3. Feb 11, 1993 · Facts. Photo from the Nobel Foundation archive. Robert W. Holley. The Nobel Prize in Physiology or Medicine 1968. Born: 28 January 1922, Urbana, IL, USA. Died: 11 February 1993, Los Gatos, CA, USA. Affiliation at the time of the award: Cornell University, Ithaca, NY, USA.

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    • Robert Holley and Nucleic Acid Structure
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    Robert Holley from Cornell University discovered a particular type of nucleic acid, transfer RNA (tRNA) and in 1965, deciphered its structure. It was the first time the structure of a biologically active nucleic acid had been established. It emerged that tRNA was the mystery molecule that Crick had proposed ten years previously in his “Adapter Hypo...

  5. Feb 14, 1993 · Robert W. Holley, a biologist who won the Nobel Prize for unraveling the genetic code of ribonucleic acid, or RNA, died Thursday at his home in Los Gatos, Calif. He was 71. The cause of death...

  6. Robert William Holley (January 28, 1922 – February 11, 1993) was an American biochemist. He shared the Nobel Prize in Physiology or Medicine in 1968 (with Har Gobind Khorana and Marshall Warren Nirenberg) for describing the structure of alanine transfer RNA, linking DNA and protein synthesis.

  7. Biochemist Robert W. Holley discovered the first nucle-otide sequence of an RNA molecule, alanine transfer RNA (tRNA). This was also, indirectly, the first nucleotide sequence of a gene. For his groundbreaking research, Holley was awarded the Nobel Prize in Physiology or Medicine in 1968, sharing the honor with Har Gobind

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